Sand Dollar (order Clypeasteroida)
InvertebrateEchinodermSeabed

The bleached test of a sand dollar (Dendraster excentricus) — this is the dead skeleton usually found on beaches, not the living animal, which is covered in dark moving spines.
Image: Björn S..., CC BY-SA 2.0, via Wikimedia Commons.
Overview
Sand dollars are flattened sea urchins — echinoderms in the order Clypeasteroida — that live on and in sandy seabeds. They are close relatives of the spiny, globe-shaped urchins found on rocky shores, redesigned for a life of burrowing in soft sediment.
The white or grey disc that people pick up on beaches is not the living animal. It is the test: the bleached internal skeleton of a dead sand dollar, stripped of its covering. The five-petalled flower pattern on it marks where tube feet emerged.
A living sand dollar is quite different — covered in a dense velvety coat of short, dark, constantly moving spines, usually purple, brown or grey, which it uses to move, burrow and pass food particles along the body.
Habitat & Range
Sand dollars live on sandy and muddy seabeds from the intertidal zone down into deeper water, in temperate and tropical seas worldwide. They are usually found in beds, sometimes at high density, either lying flat on the surface or partly buried. In areas with strong currents, young animals may swallow heavy sand grains that act as ballast to keep them from being swept away.
Diet
Sand dollars feed on fine organic particles: plankton, diatoms, algae, detritus and other material in the sediment and water. Food is caught on the spines and tube feet, moved along ciliated grooves toward the mouth on the underside, and processed by a chewing structure known as Aristotle's lantern.
Behavior
Sand dollars burrow using their spines, moving slowly across and through the sediment. In calm water they often lie flat, while in currents many species stand on edge, partly buried, to filter food from the passing flow. Dense beds are a characteristic sight in some areas. Their movement is slow and continuous, driven by the spines and tube feet rather than by any rapid locomotion.
Human Interaction & Conservation
Sand dollar tests are widely collected as beach souvenirs, and the important distinction is between a bleached, dead test and a living animal: a live sand dollar is dark and its spines move. Collecting live animals is discouraged and is regulated or prohibited in many places. Sand dollars also matter as prey for sea stars, fish and crabs, and their beds influence the structure of soft-sediment communities. This profile offers no collecting guidance; follow local rules.
Frequently Asked Questions — Sand Dollar
Is a sand dollar a shell?
What does a living sand dollar look like?
What is the flower pattern on a sand dollar?
What do sand dollars eat?
Sources and further reading
Authoritative wildlife references used for general educational context. Conservation status should always be verified against current IUCN Red List data. External links open in a new tab.
- UniversityAnimal Diversity Web — Clypeasteroida (sand dollars) — University of Michigan species account
- ReferenceSmithsonian Ocean — National Museum of Natural History — Smithsonian Institution educational ocean-science resource
- ReferenceWoRMS — World Register of Marine Species — Authoritative register of marine species names

